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Uplands Field Research Laboratory. The Balsam Woolly Aphid and Spruce-Fir Forests: A Summary of Pertinent Information for the Great Smoky Mountains National Park Interpretation Staff and the Tremont Environmental Education Center Staff In The Uplands Field Research Laboratory Volunteer-in-Parks Interpretation of Science Project. Gatlinburg, TN: Great Smoky Mountains Natural History Association , 1983.
Culver, David C.. "On Using Horn's Markov Succession Model." The American Naturalist 117, no. 4 (1981): 572-574.
Fuller, R. D.. Why Does Spruce Not Invade the High Elevation Beech Forests of the Great Smoky Mountains? In Department of Biology. Knoxville, TN: University of Tennessee, 1977.
Ramseur, George S.. Secondary Succession in the Spruce-Fir Forest of the Great Smoky Mountains National Park In Management Report. Atlanta, GA: National Park Service, Southeast Regional Office, 1976.
Norris, Daniel Howard. Bryoecology of the Appalachian Spruce-Fir Zone. Knoxville, TN: University of Tennessee, 1964.