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Comparative Studies of Macroscopic and Microscopic Features of Spermatophores of Some Plethodontid Salamanders." Copeia, no. 4 (1963): 659-669.
"Rock-Face Populations of the Mountain Salamander, Desmognathus Ochrophaeus, In North Carolina." Ecological Monographs 43, no. 1 (1973): 59-77.
"Male Secondary Sexual Characters of the Eurycea bislineata (Amphibia, Urodela, Plethodontidae) Complex in the Southern Appalachian Mountains." Journal of Herpetology 13, no. 3 (1979): 245-253.
"Hybridization between Two Species of Desmognathus (Amphibia: Caudata: Plethodontidae) in the Great Smoky Mountains." Herpetological Monographs 2 (1988): 27-39.
"Tests for Sexual Isolation Among Sympatric Salamanders of the Genus Desmognathus." Amphibia-Reptilia 11, no. 2 (1990): 147-153.
"The Courtship Behavior of the Santeetlah Dusky Salamander, Desmognathus santeetlah Tilley (Amphibia: Caudata: Plethodontidae)." Ethology 90, no. 3 (1992): 236-246.
"The Salamander Capital of the World." Smokies Life Magazine 3, no. 2 (2009): 21-27.
"Reproductive Allometry in Three Species of Dusky Salamanders." Copeia 14, no. 3 (2014): 419-427.
"Widespread Rapid Reductions in Body Size of Adult Salamanders in Response to Climate Change." Global Change Biology 20, no. 6 (2014): 1751-1759.
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