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Handley, Charles O.. "A New Flying Squirrel from the Southern Appalachian Mountains." Proceedings of the Biological Society of Washington 66 (1953): 191-194.
Pelton, Michael R.. Mammals of the Spruce-Fir Forest in Great Smoky Mountains National Park In The Southern Appalachian Spruce-Fir Ecosystem: Its Biology and Threats. Gatlinburg, TN: Uplands Field Research Laboratory, 1984.
Pfitzer, Donald W.. Recent Collections of Mammals From Great Smoky Mountains National Park. Johnson City, TN: Tennessee Academy of Science, 1951.
Singer, Francis J., David P. LaBrode, and Lorrie Sprague. Beaver Reoccupation and an Analysis of the Otter Niche in Great Smoky Mountains National Park In Research/Resource Management Report. Gatlinburg, TN: U.S. Department of Interior, National Park Service, 1981.