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Kalb, Klaus. "New or Otherwise Interesting Lichens II." Bibliotheca Lichenologica 88 (2004): 301-329.
Keller, Harold W.. "Tree Canopy Biodiversity: Student Reserach Experienced in Great Smoky Mountains National Park." Systematics and Geography of Plants 74, no. 1 (2004): 47-65.
Rhoades, Fred M.. A Review of Lichen and Bryophyte Elemental Content Literature with Reference to Pacific Northwest Species. Bellingham, WA: United States Department of Agriculture, Forest Service, 1999.
Gove, Doris. "Smokies Shelter a Diversity of Lichens." Discovering the Smokies 2, no. 1 (1999): 7-9.