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Park, Jong Seok, Christopher E. Carlton, and Michael L. Ferro. "Diversity and Taxonomic Review of Leptusa Kraatz (Coleoptera: Staphylinidae: Aleocharinae) from Great Smoky Mountains National Park, U.S.A., With Descriptions of Four New Species." \i Zootaxa\i0 , no. 2662 (2010): 1-27.\par \par }