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Sharp, Aaron J.. "Taxonomic and Ecological Studies of Eastern Tennessee Bryophytes." The American Midland Naturalist 21, no. 2 (1939): 267-354.
Stupka, Arthur. "Through the Year in the Great Smoky Mounatins National Park, Month by Month." In The Great Smokies and the Blue Ridge: The Story of the Southern Appalachians, edited by Roderick Peattie, 263-289. New York: The Vanguard Press, 1943.
Keller, Harold W.. "Tree Canopy Biodiversity: Student Reserach Experienced in Great Smoky Mountains National Park." Systematics and Geography of Plants 74, no. 1 (2004): 47-65.
Sharp, Aaron J.. "Tropical Bryophytes in the Southern Appalachians." Annals Bryologici 11 (1938): 141-144.

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