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Amphibian Monitoring in Great Smoky Mountains National Park In Planning for the Seventh Generation: The Upper Great Lakes Region 25th Annual Natural Areas Association Conference. Mackinac Island, MI: Natural Areas Association, 1998.
Frequency of Hybrids between Introduced and Native Populations of the Salamander Plethodon jordani in Their First Generation of Sympatry." Herpetologica 54, no. 2 (1998): 143-153.
"The Dynamics of Two Hybrid Zones in Appalachian Salamanders of hte Genus Plethodon." Evolution 46, no. 4 (1992): 930-038.
"Hybridization between Two Species of Desmognathus (Amphibia: Caudata: Plethodontidae) in the Great Smoky Mountains." Herpetological Monographs 2 (1988): 27-39.
"Species Discrimination between Hybridizing and Non-Hybridizing Terrestrial Salamanders." Copeia 1987, no. 4 (1987): 924-931.
"Southern Appalachian Salamanders: An Indexed Bibliography In Research/Resources Management Report. Gatlinburg, TN: U.S. Department of the Interior, National Park Service, 1985.
Herpetofauna of the Spruce-Fir Ecosystem in the Southern Appalachian Mountain Regions, With Emphasis on the Great Smoky Mountains National Park In The Southern Appalachian Spruce-Fir Ecosystem: Its Biology and Threats. Gatlinburg, TN: Uplands Field Research Laboratory, 1984.
A New Species of Desmognathus (Amphibia: Caudata: Plethodontidae) from the Southern Appalachian Mountains In Occasional Papers of the Museum of Zoology. Vol. 695. Ann Arbor, MI: Musuem of Zoology, University of Michigan, 1981.
Amphibians and Reptiles of the Carolinas and Virginia. Chapel Hill, NC: University of North Carolina Press, 1980.
Evolution Under Interspecific Competition: Field Experiments on Terrestrial Salamanders." Evolution 34, no. 3 (1980): 409-420.
"The Experimental Test of an Analysis of Field Distributions: Competition In Terrestrial Salamanders." Ecology 61, no. 4 (1980): 817-826.
"Male Secondary Sexual Characters of the Eurycea bislineata (Amphibia, Urodela, Plethodontidae) Complex in the Southern Appalachian Mountains." Journal of Herpetology 13, no. 3 (1979): 245-253.
"The Distribution of the Fungus, Basidiobolus Ranarum Eidam, in Fish, Amphibians, and Reptiles of the Southern Appalachian Region." Transactions of the Kansas Academy of Science 80, no. 1-2 (1977): 75-77.
"The Southern Appalachians In The American Wilderness. New York: Time-Life Books, 1975.
Experimental Study of Batesian Mimicry in the Salamanders Plethodon Jordani and Desmognathus ochrophaeus." The American Midland Naturalist 90, no. 1 (1973): 38-46.
"Normal Blood Values in Some Plethodontid Salamanders." Herpetologica 29, no. 2 (1973): 184-188.
"Rock-Face Populations of the Mountain Salamander, Desmognathus Ochrophaeus, In North Carolina." Ecological Monographs 43, no. 1 (1973): 59-77.
"Altitudinal Distribution of the Protozoan Cepedietta michiganesis in the Salamanders Plethodon glutinosus and Plethodon jordani." American Midland Naturalist 83, no. 2 (1970): 393-403.
"Homing Behavior of the Red-Checked Salamander, Plethodon Jorani." Animal Behavior 17, no. 1 (1969): 25-39.
"Amphibians and Reptiles of the Great Smoky Mountains National Park. Knoxville, TN: University of Tennessee, 1967.
The Desmognathine Salamanders of the Great Smoky Mountains National Park." Journal of the Ohio Herpetological Society 5, no. 3 (1966): 63-72.
"The Appalachians. Boston: Houghton Mifflin, 1965.
Herpetological Records from the Great Smoky Mountains." Herpetologica 21, no. 2 (1965): 148-150.
"Further Notes on Mimicry in Salamanders." Herpetologica 17, no. 1 (1961): 63-64.
"Great Smoky Mountains National Park In Natural History Handbook. Vol. 5. Washington, D.C.: U.S. Government Printing Office, 1961.