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Land Mollusca of Great Smoky Mountains National Park. Great Smoky Mountains National Park Research Library, Undated.
Records and Descriptions of North American Crane-Flies (Diptera) Part II. Tipuloidea of Mountainous Western North Carolina." The American Midland Naturalist 26, no. 2 (1941): 281-319.
" A New Species of Polygra from the Great Smoky Mountains." Nautilus 51, no. 4 (1938): 135-137.
"Evolution of the (Coleoptera) Carabidae in the Southern Appalachians." In The Distributional History of the Biota of the Southern Appalachians. Part I: Invertebrates, 67-92. Virginia Polytechnic Institute and State University. Research Division. Monographs. Blacksburg, VA: Virginia Polytechnic Institute, 1969.
"The North American Pterostichus of the Subgenus Cylindrocharis Casey (Coleoptera, Carabidae)." American Museum Novitates, no. 2445 (1971): 1-14.
"New Spiders from the Great Smoky Mountain National Park." Annals of the Entomological Society of America 38, no. 1 (1945): 70-76.
"Ramazzottius belubellus, a new species of Tardigrada (Eutardigrada: Parachela: Hypsibiidae) from the Great Smoky Mountains National Park (North Carolina, U.S.A.)." Proceedings of the Biological Society of Washington 124, no. 1 (2011): 23-27.
"Macrobiotus (Eutardigrada, Macrobiotidae) from the Great Smoky Mountains National Park, Tennessee/North Carolina, USA (North America): Two New Species and Six New Records." Zootaxa, no. 2022 (2009): 45-57.
"Doryphoribius smokiensis, A New Species of Eutardigrada (hypsibiidae) from the Great Smoky Mountains National Park, TN, USA (North America)." Zootaxa, no. 1646 (2007): 59-65.
"Two New Genera and Species of Stenocranine Planthoppers (Hemiptera : Delphacidae) from North America." Entomological News 116, no. 5 (2005): 291-303.
"The Great Smoky Mountains Bat Blitz of 2002." Bat Research News 43, no. 4 (2002): 135-136.
"The Natural History and Taxonomy of Cicurina Bryantae Exline (Araneae, Agelenidae)." Journal of Arachnology 13, no. 1 (1985): 87-96.
"Moruliua delicata, New Species from Great Smoky Mountains National Park and Redescriptions of M-callowayia Wray and M-crassa Christiansen & Bellinger (Collembola : Neanuridae)." Proceedings of the Biological Society of Washington 119, no. 4 (2006): 540-556.
"Trichodorus elefjohnsoni n. sp.(Nemata: Trichodoridae) from Undisturbed Appalachian Forest." Journal of Nematology 24, no. 1 (1992): 78-83.
"Furculanurida langdoni n. sp (Collembola : Neanuridae), a Nearctic Member of a Gondwanan genus." Proceedings of the Biological Society of Washington 120, no. 3 (2007): 320-326.
"Redescription of Cosberella conatoa Wray (Collembola : Hypogastruridae) and the Description of C-lamaralexanderi." Proceedings of the Biological Society of Washington 119, no. 2 (2006): 269-278.
"Trap Success of Striped Skunks (Mephitis mephitis) in Great Smoky Mountains National Park." Journal of the Tennessee Academy of Science 79, no. 4 (2004): 91-93.
"Ferns of North Carolina. Durham, NC: Duke University Press, 1934.
Wild Hogs in the United States -- Origin and Nomenclature In Research & Management of Wild Hog Populations: Proceedings of a Symposium. Georgetown, South Carolina: Belle W. Baruch Forest Science Institute of Clemson University, 1977.
Bats of Great Smoky Mountains National Park and Nantahala National Forest." Bat Research News 40, no. 4 (1999): 164.
"Lepidoptera TWIG Branches Out." ATBI Quarterly 2, no. 4 (2001): 7.
"Discover Life in America." Smoky Mountain Living 3, no. 4 (2003): 116-117.
"Mining Beetle Diversity in the Smokies." ATBI Quarterly 2, no. 4 (2001): 3.
"Smoky Mountain Reptiles." ATBI Quarterly 2, no. 4 (2001): 6.
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