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Bertolani, Roberto, Paul J. Bartels, Roberto Guidetti, Michele Cesari, and Diane R. Nelson. "Aquatic Tardigrades in the Great Smoky Mountains National Park, North Carolina and Tennessee, U.S.A., with the Description of a New Species of Thulinius (Tardigrada, Isohypsibiidae)." Zootaxa 3764, no. 5 (2014): 524-536.
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Davison, Paul G.. "Feeding Water Bears: A Simple Activity to Connect the Public with Microorganisms." Southeastern Biology 53, no. 1 (2006): 32-34.
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Bartels, Paul J., Diane R. Nelson, Lukasz Kaczmarek, and Lukasz Michalczyk. "The Genus Milnesium (Tardigrada: Eutardigrada: Milnesiidae) in the Great Smoky Mountains National Park." Zootaxa 3826, no. 3 (2014).
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Bartels, Paul J.. "Have You Seen This Creature?" ATBI Quarterly 2, no. 2 (2001): 1.
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Ray, M. W., H. F. Heatwole, and Thomas R. Wentworth. Limno-Terrestrial Tardigrades of North Carolina: Initial Findings of a Study Along an East-West Transect. 3rd ed. Vol. 55. Burlington, NC: Association of Southeastern Biologists, Inc, 2008.
Bartels, Paul J.. ""Little known" Water Bears." Southeastern Biology 53, no. 1 (2006): 30-32.
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Bartels, Paul J., Giovanni Pilato, Oscar Lisi, and Diane R. Nelson. "Macrobiotus (Eutardigrada, Macrobiotidae) from the Great Smoky Mountains National Park, Tennessee/North Carolina, USA (North America): Two New Species and Six New Records." Zootaxa, no. 2022 (2009): 45-57.
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Bartels, Paul J., Diane R. Nelson, Lukasz Kaczmarek, and Lukasz Michalczyk. "Ramazzottius belubellus, a new species of Tardigrada (Eutardigrada: Parachela: Hypsibiidae) from the Great Smoky Mountains National Park (North Carolina, U.S.A.)." Proceedings of the Biological Society of Washington 124, no. 1 (2011): 23-27.
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Nelson, Diane R., and Paul J. Bartels. ""Smoky Bears" - Tardigrades of Great Smoky Mountains National Park." Southeastern Naturalist 6, no. Special Issue 1 (2007): 229-238.
Nelson, Diane R., and Paul J. Bartels. "Species Richness of Soil and Leaf Litter Tardigrades in the Great Smoky Mountains National Park (North Carolina/Tennessee, USA)." Journal of Limnology 72, no. s1 (2013): 144-151.